Name change chain

Red Barchetta (March 19, 2011) -> A New World Record (November 17, 2012) -> Lucina (March 18, 2013) -> Dean Winchester (May 20, 2013) -> Boy George (November 01, 2014) -> Palkia (March 05, 2015) -> Nino (April 05, 2015) -> Chloe Beale (May 09, 2015) -> Kirstie Maldonado (March 22, 2016) -> Lauren Mayberry (June 27, 2016) -> Araquanid (November 23, 2016) -> Miles Edgeworth (January 03, 2017) -> Godot (March 18, 2017)
 
Roserade → Tsareena → Lilligant
 
2013
Fawful (March 2013)

Lord Spongeful (May 2013)

King Antasma (June 2013)

His Excellency Lord Fawfulthegreat64 of the Beanbean Realm (July 2013)

Fawfulthegreat64 (August 2013)

Squidward Tentacles (September 2013)

Count FTG64 (October 2013)

Dimentio64 (November 2013)

FawfulTheSnowman (December 2013)

2014
Decade of Luigi (January 2014)

Count Bleck (February 2014)

Fawfulthegreat64 (March 2014)

Dreamy Fawful (May 2014)

Too Bad. Waluigi Time. (June 2014)

That One Weird Dude Who Plays Mario And Eats Burgers All Day (July 2014)

Paper Mario (August 2014)

Super Smash Bros. for Nintendo 3DS (September 2014)

King Antasma (October 2014)

Fawfulthegreat64 (November 2014)

Snawfulthemerry25 (December 2014)

2015
Stanley the Troll (January 2015)

Wittgenstein (February 2015)

Fawful (March 2015)

Fawfulthegreat64 (April 2015 - January 2017)

2017
Fresh Bowser (January-February 2017)

Air Conditioner (March 2017)

greatdimentio (April 2017)
 
MegaDigga3
HydroGuy3503
Man Without Hat
Tumble
Police Squad!
Some others (Various 2013-2014)
Jonny Davis (August 2014-September 2015)
JG Wentworth (September 2015)
Cahoots Is Spelled With A C (October 2015)
Cat Gray (November-December 2015)
Mary Berry (January 2016)
Roy Koopa (February 2016)
Laverne Todd (March 2016)
Charley Dietz (April-May 2016)
F-777 (May-June 12? 2016)
Bob Eubanks (June 12?-September 19, 2016)
Jim Perry (September 19-October 25, 2016)
Gene Rayburn (October 25, 2016-November 25, 2016)
Michael Palin (November 25, 2016-December 25, 2016)
Bob McKenzie (December 25, 2016-January 27, 2017)
Rick Lax (January 27, 2016-February 26, 2017)
Bob Eubanks (February 26, 2017-March 26, 2017)
SpaceHamster (March 26, 2017-)
 
Red Barchetta (March 19, 2011) -> A New World Record (November 17, 2012) -> Lucina (March 18, 2013) -> Dean Winchester (May 20, 2013) -> Boy George (November 01, 2014) -> Palkia (March 05, 2015) -> Nino (April 05, 2015) -> Chloe Beale (May 09, 2015) -> Kirstie Maldonado (March 22, 2016) -> Lauren Mayberry (June 27, 2016) -> Araquanid (November 23, 2016) -> Miles Edgeworth (January 03, 2017) -> Godot (March 18, 2017) -> (April 18, 2017)

legendary
 
Roserade → Tsareena → Lilligant → Taranza
 
November 22, 2016 - March 22, 2017: Luigi 64DD
March 22, 2017 - April 22, 2017: Smiley Man
April 22, 2017 - present: Luigi 64DD

I apparently got my first name change exactly 4 months after I joined without realizing it. What a coincidence!
 
Pyro Guy --> The Pyro Guy --> The Universe --> The Pyro Guy --> Cory Baxter --> Spooky the slightly frightening Shy Guy --> The Pyro Guy --> Turtonator --> The Pyro Guy --> Chef Kawasaki
 
time for update

Season 1
​Kingbowser99 (15 Jan 09 - 09 Aug 12) 1302 days
Bane (09 Aug 12 - 28 Sep 12) 50 Days
Radagast the Brown (28 Sep 12 - 03 Dec 12) 66 days
Santagast the Merry (03 Dec 12 - 26 Dec 12) 23 days
Radagast the Brown (26 Dec 12 - 25 Jan 13) 30 days
Sochies (25 Jan 13 - 01 Mar 13) 35 days
The End (01 Mar 13 - 18 Apr 13) 48 days

Season 2
Dozla (18 Apr 13 - 19 May 13) 31 days
Neptune 2.0 (19 May 13 - 20 Jun 13) 32 days
Joey Tribbiani (20 Jun 13 - 01 Aug 13) 42 days
Kingbowser99 (01 Aug 13 - 03 Sep 13) 33 days
Radagast the Brown (03 Sep 13 - 03 Nov 13) 61 days
Delelelele Whoop (03 Nov 13 - 03 Dec 13) 30 days
War Doctor (03 Dec 13 - 25 Mar 14) 112 days

Season 3
Mrs. Doubtfire (25 Mar 14 - 29 Apr 14) 35 days
Иeptune (29 Apr 14 - 05 Aug 14) 98 days
Priam (05 Aug 14 - 24 Sep 14) 50 days
Mycroft Holmes (24 Sep 14 - 25 Dec 14) 92 days
Hoxton (25 Dec 14 - 27 Mar 15) 92 days
Matthew Bellamy (27 Mar 15 - 28 Apr 15) 32 days

Season 4
Ultron (28 Apr 15 - 05 Mar 16) 313 Days
Leonardo DiCaprio's Oscar (05 Mar 16 - 7 Apr 16) 32 Days
General Neptune (7 Apr 16 - 25 Jun 16) 78 Days
JonTron (25 Jun 16 - 27 Jul 16) 32 Days
Jeff Goldblum (27 Jul 16 - 18 Nov 16) 114 Days
Decidueye (18 Nov 16 - 3 Apr 17) 136 Days

Season 5
Brian Butterfield (3 Apr 17)

if you're curious what these seasons mean, i consider each of these usernames to be different milestones of my personality. S1 is a complete dickish, immature Kingbowser99. S2 is a much more mature Neptune, but someone who is incredibly arrogant and controlling. S3 is someone who is almost there, a guy who realises that his head-on approach is only causing anger and sadness, so he adopts a PG personality, still with the whole "immortality and wizardry w00t" arrogance. S4 is a much more down to earth chap, who knows how to speak for himself without sounding like a dick. He knows his friends, and tries to better himself by building friendships with people that he wouldn't normally be considered close to. S5 is the lovable comedy idiot phase, wouldn't pay too much attention to this guy.
 
2013
July 31: thatmariochick
October 26: EmeraldDiamond

2014
April 11: Astronomical
May 18: Jolteon
June 18: Princess Rosalina
July 17: Diantha
August 12: Stargazing
September 12: Rosalina and Luma
October 13: Stargazing
November 11: Vaporeon
December 12: Snowzomi Mistletoujou

2015
January 12: Nozomi Toujou
February 13: Stargazing
March 19: Nozomi Toujou
April 18: Stargazing
May 18: May
June 16: Nozomi Toujou
July 16: Stargazing
August 15: Ness
September 15: Stargazing
November 8: Nozomi Toujou

2016
January 23: Stargazing
February 24: Ness
March 23: Nozomi Toujou
May 21: Kanan Matsuura
July 31: Nozomi Toujou
August 29: Stargazing
October 16: Kanan Matsuura
November 19: Lillie
December 22: Stargazing

2017
February 7: Chara
March 6: Stargazing
April 22: Riko Sakurauchi